Department of Archeology, University of Durham, Durham, United Kingdom
Angela R. Perri
Australian Center for Ancient DNA, School of Biological Sciences, University of Adelaide, Adelaide, South Australia, Australia
Kieren J. Mitchell, Graham Gower, Holly Heiniger and Alexander T. Salis
Department of Ecology and Evolutionary Biology, University of California, Los Angeles, California, USA
Alice Mouton, Colin Shew, Blaire Van Valkenburgh and Robert K. Wayne
School of Biological and Chemical Sciences, Queen Mary University of London, London, United Kingdom
Sandra Álvarez-Carretero, Alberto Carmagnini, Mario dos Reis and Laurent AF Frantz
Department of Archeology, Classics and Egyptology, University of Liverpool, Liverpool, UK
Ardern Hulme-Beaman and Keith Dobney
School of Natural Sciences and Psychology, John Moores University of Liverpool, Liverpool, UK
Ardern Hulme-Beaman
The Palaeogenomics & Bio-Archeology Research Network, Research Laboratory for Archeology and History of Art, University of Oxford, Oxford, United Kingdom
James Haile, Alexandra Jamieson, Audrey T. Lin, Anna Linderholm and Greger Larson
Department of Anatomy, Des Moines University, Des Moines, IA, USA
Julie meachen
Department of Zoology, University of Oxford, Oxford, United Kingdom
Audrey T. Lin
Department of Anthropology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA
Audrey T. Lin
Center of Excellence in Paleontology and Department of Geosciences, East Tennessee State University, Johnson City, TN, USA
Blaine W. Schubert
Department of Archeology, University of Exeter, Exeter, UK
Carly Ameen
Institute of Archeology, Russian Academy of Sciences, Moscow, Russia
Ekaterina E. Antipina
ARAID Foundation, University Institute for Research in Environmental Sciences (IUCA) – Aragosaurus Group, University of Zaragoza, Zaragoza, Spain
Father Bover
Department of Earth Sciences, Natural History Museum, London, United Kingdom
Selina brace
Evolutionary Genomics Section, GLOBE Institute, University of Copenhagen, Copenhagen, Denmark
Christian Carøe, Jose A. Samaniego Castruita, Shyam Gopalakrishnan, Matthew J. Collins, Mikkel-Holger S. Sinding and M. Thomas P. Gilbert
Applied Paleoscience, Bothell, WA, United States
James C. Chatters
Department of Archeology, University of Sydney, Sydney, New South Wales, Australia
Keith Dobney
Department of Archeology, University of Aberdeen, Aberdeen, United Kingdom
Keith Dobney
Department of Archeology, Simon Fraser University, Burnaby, Canada
Keith Dobney
Institute of Evolutionary Sciences – Montpellier, CNRS, University of Montpellier, IRD, EPHE, Montpellier, France
Allowen evin
Evolution & Biological Diversity Laboratory, UPS / CNRS / IRD, Paul Sabatier University, Toulouse, France
Philippe gaubert
Australian Museum Research Institute, Australian Museum, Sydney, New South Wales, Australia
Kristofer M. Helgen
Department of Ecology and Evolutionary Biology, University of California at Santa Cruz, Santa Cruz, California, USA
Josh Kapp, Nedda F. Saremi and Beth Shapiro
Institute of Plant and Animal Ecology, Urals Branch of the Russian Academy of Sciences, Yekaterinburg, Russia
Pavel A. Kosintsev
Federal University of the Urals, Yekaterinburg, Russia
Pavel A. Kosintsev
Department of Anthropology, Texas A&M University, College Station, TX, USA
Anna linderholm
Center for Evolution and Medicine, Arizona State University, Tempe, AZ, USA
Andrew T. Ozga and Anne C. Stone
School of Human Evolution and Social Change, Arizona State University, Tempe, AZ, USA
Andrew T. Ozga, Katherine Skerry and Anne C. Stone
Halmos College of Arts and Sciences, Nova Southeastern University, Fort Lauderdale, FL, USA
Andrew T. Ozga
Department of Archeology, University of York, York, United Kingdom
Samantha presslee
Institute of Animal Systematics and Ecology, Siberian branch of the Russian Academy of Sciences, Novosibirsk, Russia
Dmitry E. Taranenko
Idaho Museum of Natural History, Idaho State University, Pocatello, ID, USA
Mary thompson
Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia
Mikhail V. Sablin
Sobolev Institute of Geology and Mineralogy, Siberian branch of the Russian Academy of Sciences, Novosibirsk, Russia
Yaroslav V. Kuzmin
Tomsk State University, Tomsk, Russia
Yaroslav V. Kuzmin
McDonald’s Institute for Archaeological Research, University of Cambridge, Cambridge, UK
Matthew J. Collins
Greenland Natural Resources Institute, Nuuk, Greenland
Mikkel-Holger S. Sinding
NTNU University Museum, Trondheim, Norway
Mr. Thomas P. Gilbert
Institute of Human Origins, Arizona State University, Tempe, AZ, USA
Anne C. Stone
Howard Hughes Medical Institute, University of California Santa Cruz, Santa Cruz, California, USA
Beth shapiro
South Australian Museum, Adelaide, South Australia, Australia
Alan cooper
Paleogenomics Group, Department of Veterinary Sciences, Ludwig Maximilian University, Munich, Germany
Laurent AF Frantz
ARP, KJM, AM, RKW, GL, LAFF and A. Cooper designed the project and designed the research; ARP and KJM coordinated the sample collection efforts with input from RKW, GL, LAFF and A. Cooper; ARP, KJM, AH-B., JM, CA, JCC, AE, PG, JK, AL, ATO, SP, BWS, MT, MJC, M.-HSS, MTPG, ACS, BS, BVV, RWK and A. Cooper provided and / or collected samples; ARP, KJM, RKW, AM, CS, JH, AJ, ATS, PB and HH conducted the genomic lab work; KJM, AM, GG, GL, LAFF and A. Cooper performed the analyzes of the genomic data; SA-C., AH-B., JM, CA, KMH and AE performed the morphological analyzes; ARP, KJM, AM, SA-C., BVV, KMH, RKW, GL, LAFF and A. Cooper wrote the article with contributions from all other authors.